The Armstrong-Rauk Oxidative Damage to Proteins Research Group

(last update April 2, 2002)

Research Projects:

Selected Publications

161. P. Brunelle and A. Rauk, Specific Recognition of Gly29 and Gly33 by Met35 in a Model of b-Sheet Ab: An ONIOM Study, J. Alzheimer's Disease, in press (accepted 2002/03/04) - Abstract

160. D. L. Reid, G. V. Shustov, D. A. Armstrong, A. Rauk, M. N. Schuchmann, M. S. Akhlaq, and C. von Sonntag, H-Abstraction from Thiols by C-Centered Radicals. An Experimental and Theoretical Study, Phys. Chem. Chem. Phys. in press (accepted (2001/11/05.) - Abstract

157. A. Rauk, D. A. Armstrong, and J. Berges, Glutathione Radical: Intramolecular H Abstraction by the Thiyl Radical, Can. J. Chem.,79, 405-417 (2001) - Abstract

156. A. Rauk, D. A. Armstrong, and D. P. Fairlie, Is Oxidative Damage By Beta Amyloid and Prion Peptides Mediated by Hydrogen Atom Transfer from Glycine Alpha-Carbon to Methionine Sulfur within Beta-Sheets?, J. Am. Chem. Soc. 122, 9761-9767 (2000) - Abstract

153. A. Rauk and D. A. Armstrong, Influence of beta-Sheet Structure on the Susceptibility of Proteins to Backbone Oxidative Damage: Preference for alpha-C-Centered Radical Formation at Glycine in Antiparallel beta-Sheets, J. Am. Chem. Soc, in press (accepted 2000/02/02) - Abstract

150.  A. Rauk, D. Yu, J. Taylor, G. V. Shustov, D. A. Block, and D. A. Armstrong, A Comparison of the alpha-C-H Bond Enthalpies of Amino Acid Residues in a Protein Model Environment, Biochemistry, 38, 9089-9096 (1999) - Abstract

146. G. V. Shustov and A. Rauk, Dioxirane Oxidation of Nitrosamines. An Ab Initio Study,  Can. J.  Chem.,  77, 74-85 (1999).

143. D. A. Block, D. Yu, D. A. Armstrong, and A. Rauk, On the Influence of Secondary Structure on the a-C-H Bond Dissociation Energy of Proline Residues in Proteins: a Theoretical Study,  Can. J. Chem., 76, 1042-1049 (1998)..

142. M. Jonsson, D. D. M. Wayner, D. A. Armstrong, D. Yu, and A. Rauk, On the Thermodynamics of Peptide Oxidation, J. Chem. Soc. Perkin II, 1967-1972 (1998).

141. G. V. Shustov and A. Rauk, A Theoretical Study of Oxidation of CH Bonds in Homo- and Heterosubstituted Alkanes by Dioxirane as a Model of the Dioxirane Oxidation of Peptides, J. Org. Chem.63, 5413-5422 (1998).

140. D. A. Armstrong, D. Yu, and A. Rauk,  Oxidative Damage to Cysteine in Proteins: an Ab Initio Study of the Radical Structures, C-H, S-H, and C-C Bond Dissociation Energies, and Transition Structures for H Abstraction by Thiyl Radicals, J. Am. Chem. Soc. 120, 8848-8855 (1998). - Abstract

135. D. D. M. Wayner, K. B. Clark, A. Rauk, D. Yu, and D. A. Armstrong, C-H Bond Dissociation Energies of Alkyl Amines, J. Am. Chem. Soc., 119, 8925-8932 (1997).

131. D. A. Armstrong, D. Yu, and A. Rauk,  Gas Phase and Aqueous Thermochemistry of Hydrazine and Related Radicals and the Energy Profiles of Reactions with H. and OH.: An Ab Initio Study, J. Phys. Chem. 101, 4761-4769 (1997).

128. A. Rauk, D. Yu, and D. A. Armstrong, Toward Site Specificity of Oxidative Damage in Proteins: C-H and C-C Bond Dissociation Energies and the Reduction Potentials of the Radicals of Alanine, Serine, and Threonine - an Ab Initio Study, J. Am. Chem. Soc. 119, 208-217 (1997). - Abstract

121. D. A. Armstrong, D. Yu, and A. Rauk, Oxidative Damage to the Glycyl a-Carbon in Proteins: an Ab Initio Study of the C-H Bond Dissociation Energy and the Reduction Potential of the C-Centered Radical, Can. J. Chem., 76, 1192-1199 (1996).

117. D. Yu, D. A. Armstrong and A. Rauk, The Structures and Relative Energies of Formamide (H2NCHO) and Radical Ions H2NCHO.+, H2NCOH.+, and H3NCO.+, Chem. Phys. 202, 243 (1996).

109. A. Rauk, D. Yu, P. Borowski, and B. Roos, CASSCF, CASSPT2, and MRCI Investigations of Formyloxyl Radical (HCOO.), Chemical Physics, 197, 73-80 (1995) - reprints not ordered.

105. D. Yu, A. Rauk, and D. A. Armstrong, The Solution Thermochemistry of the Radicals of Glycine, J. Chem. Soc. Perkin 2, 553 - 560 (1995).

104. D. Yu, A. Rauk, and D. A. Armstrong, The Radicals and Ions of Glycine: An ab initio Study of the Structures and Gas Phase Thermochemistry , J. Am. Chem. Soc. 117, 1789 - 1796 (1995).

99. D. Yu, A. Rauk, and D. A. Armstrong, The Radicals and Ions of Formic and Acetic Acids Acids: An ab initio Study of the Structures and Gas and Solution Phase Thermochemistry, J. Chem. Soc. Perkin 2, 2207 - 2215 (1994).

97. A. Rauk, D. Yu, and D. A. Armstrong, Carboxyl Free Radicals: Formyl and Acetyl Revisited, J. Am. Chem. Soc., 116, 8222-8228 (1994).

93. D. Yu, A. Rauk, and D. A. Armstrong , Gas and solution phase thermochemistry and transition energies of NH2., NH3+., and their aquo complexes: An ab initio study. Can. J. Chem., 72, 471-483 (1994)

92. A. Rauk, D. A. Armstrong, and D. Yu, The Lifetime of Gas Phase CO2.- and N2O.- Calculated from the Transition Probability of the Autodetachment Process A- - A + e-. Int. J. Chem. Kinet., 26, 7 (1994)



161.  Specific Recognition of Gly29 and Gly33 by Met35 in a Model of b-Sheet Ab: An ONIOM Study,

Abstract: The Radical Model of  Alzheimer’s Disease (AD) is presented in some detail.  The model provides a unified picture for the role of the amyloid beta peptide (Ab), Met35, copper ions, oxygen, beta sheet secondary structure, and the generation of hydrogen peroxide, in mediating oxidative stress in AD.  It predicts a role for glycyl radicals as long-lived species which can transport the damage into cell membranes and initiate  lipid peroxidation.  Previous work has established the thermodynamic and kinetic viability of most of the steps.  In the present work, QM/MM and Amber calculations reveal that self assembly of antiparallel b-sheet which brings Met35 into the required close proximity to a glycine residue is more likely if the residue is Gly29 or Gly33, than any of the other four glycine residues of Ab.
 - Return to Selected Publications



160.  H-Abstraction from Thiols by C-Centered Radicals. An Experimental and Theoretical Study,

Abstract:  The hydrogen atom abstraction by a series of carbon-centered radicals from methanethiol is examined in the gas phase and in aqueous solution using quantum mechanical calculations.  The gas phase reactions are modeled at the ab initio B3LYP/6311+G(d,p) level, coupled with an empirical correction to the enthalpy of reaction and activation.  The solvent effects are evaluated by two different continuum models (SCIPCM, CPCM), coupled with a novel approach to the calculation of the solution phase entropy.  The reaction is discussed in terms of the charge and spin polarization in the transition state, as determined by AIM analysis, and in terms of orbital interaction theory.  Rate constants, calculated by transition state theory are in good agreement with the available experimental data.- Return to Selected Publications



157. Glutathione Radical: Intramolecular H Abstraction by the Thiyl Radical

Abstract:  Ab initio computations (B3LYP/6-31G(D)) were used to predict transition structures and energies of activation for intramolecular H atom transfer to a thiyl radical (RS.) from the aC-H bonds of glutathione 1 and from the model compounds, N-formylcysteinylglycine 2 and N-(2-thioethanyl)-g-glutamine 3. For each compound, transition structures were located by in vacuo calculations on the neutral non-zwitterionic system.   Thermodynamic functions derived at the same level and single point calculations at the B3LYP/6-311+G(3df,2p) level, were used to derive free energies of activation (DG?) and reaction (DGo).  For abstraction of the aC-H(Gly) by the thiyl radical in the gas phase, DG? = 134 kJ mol-1 if the amide link to Gly is in the more stable (Z)-configuration and DG? = 52 kJ mol-1, if it is in the less stable (E)-configuration.  The isomerization of the amide group requires about 95 kJ mol-1.  Previous studies had indicated that for intramolecular reaction of the thiyl radical at aC-H(Cys),  DG? = 110 kJ mol-1.  The lowest energy pathway for intramolecular H transfer to the thiyl radical is from aC-H(Gln), DG? =  37 - 42 kJ mol-1, and corresponds rather well with experimental results in solution, DG? =  43 kJ mol-1.  The calculated free energy change for the equilibrium between thiyl and aC forms of the glutathione radical, DGo =  -54 kJ mol-1.  The value estimated from experimental data is DGo =  -37 kJ mol-1.  The agreement between the energies from theory in the gas phase and experiment in solution suggests that the free energies of solvation of reactant thiyl radical, transition structures for H abstraction, and the product aC-centred radical, are very similar.  The effects of solution were estimated by two continuum models, SCIPCM and COSMO.  The SCIPCM model yields results very similar to the gas phase, predicting a modest lowering of the activation free energy.  The results from the COSMO method were inconclusive as to whether a rate enhancement or decrease could be expected.- Return to Selected Publications



156. Is Oxidative Damage By Beta Amyloid and Prion Peptides Mediated by Hydrogen Atom Transfer from Glycine Alpha-Carbon to Methionine Sulfur within Beta-Sheets?

Abstract: Methionine in glycine rich regions of both beta amyloid peptide and prion peptide is thought to be crucial to their neurotoxic properties.  We postulate here a role for methionine in the propagation of oxidative damage.  The S-H bond dissociation enthalpies, BDE(S-H)s, of dimethylsulfonium ion (CH3)2SH+, and a S-protonated methionine residue of a polypeptide strand are estimated to be 351 kJ mol-1 and 326 - 331 kJ mol-1 , respectively, by the application of calculations at the B3LYP level with large basis sets.  These species are direct products of H atom abstraction by radical cations of sulfides.  The reactions between a glycine residue and the radical cations of (CH3)2S and Met were investigated, and the transition structures for H atom transfer located. The results suggest that it is thermodynamically feasible for the S-ionized form of Met to cause oxidative damage at the aC-H site of almost any amino acid residue of a nearby polypeptide strand (BDE(aC-H) = 330 - 360 kJ mol-1), or to nearby lipids with a bis(allylic) methylene group (BDE(C-H) = 335 kJ mol-1).  However, a key observation is that when the Met residue is incorporated into an antiparallel b-sheet, only a Gly residue is exposed and susceptible to oxidation at the aC-H site.  Furthermore, the Gly must lie on a different strand of the b-sheet to that containing Met, and must be part of a (5,5), rather than a (3,3) cycle. The same considerations apply to the methyl-deprotonated form of the sulfide radical cation but not the methylene-deprotonated form.  These findings suggest a possible mechanism for generating and propagating oxidative damage via a Met residue of the Ab peptide of Alzheimer's Disease and of the prion peptide of Creutzfeldt-Jakob Disease.  To our knowledge, this is the first proposed mechanism that accounts for the radical damage in either of these diseases and requires peptide b-sheets, and amino acids, methionine and glycine.- Return to Selected Publications



153. Influence of beta-Sheet Structure on the Susceptibility of Proteins to Backbone Oxidative Damage: Preference for alpha-C-Centered Radical Formation at Glycine in Antiparallel beta-Sheets

Abstract: Ab initio calculations at the B3LYP/6-31G(d) level of theory were carried out on selected cyclic hydrogen bonded dimers of glycine and alanine as models for b-sheets, and on the aC-centered radicals derived from them.  The structures mirrored the cycles found in the H-bonded network of parallel and antiparallel b-sheet secondary structure, and were optimized both with and without enforcement of constraints on the F,Y torsion angles.  Transition structures for the migration of an H atom from an aC site to another aC site or to an S atom were located.  It was found that the presence of a hydrogen bonded strand of a b-sheet has little effect on the aC-H bond dissociation enthalpy (BDE) of glycine, but raises the BDE of other residues by a significant amount.  The parallel b-sheet structure and F,Y angles lead to a significant increase in BDE relative to the random coil structure, due to loss of captodative stabilization.  The antiparallel b-sheet structure and F,Y angles do not lead to a significant increase in BDE.  All residues incorporated in b-sheet secondary structure, with the exception of glycine, are protected from oxidative damage because the aC-H bond is internal to the sheet and inaccessible to oxidizing radicals.  Glycine is susceptible to oxidative damage because it has a second aC-H bond which is exposed.  Among residues in secondary structures, only glycine is susceptible to damage by weak oxidants such as thiyl radicals and superoxide, provided it is in an antiparallel b-sheet.  Radical damage may propagate readily from one strand to another above the b-sheet, but not within the b-sheet. b-Sheet structure narrows the difference between the glycyl aC-H BDE and S-H BDE and facilitates interstrand H atom transfer between the glycyl aC site and the S atom of cysteine.- Return to Selected Publications



150. Effects of Structure on aC-H Bond Enthalpies of Amino Acid Residues: Relevance to H Transfers in Enzyme Mechanisms and in Protein Oxidation

Abstract: The bond dissociation enthalpies (BDE) of all of the amino acid residues, modelled by HC(O)NHCH(R)C(O)NH2 (PH(Res)) were determined at the B3LYP/6-31G(D) level, coupled with isodesmic reactions.  The results for neutral side chains with f, y angles ~180o,~180o in ascending order, to an expected accuracy of " 10 kJ mol-1, are: Asn 326; Cystine 330; Asp 332; Gln 334; Trp 337; Arg  340; Lys 340; Met 343; His 344; Phe 344; Tyr 344; Leu 344; Ala 345; Cys 346; Ser 349; Gly 350; Ile 351; Val 352; Glu 354; Thr 357; Pro-cis 358; Pro-trans 369.  These BDEs are smaller than those of typical secondary or tertiary C-H bonds due to the phenomenon of captodative stabilisation. The stabilisation is reduced by changes in the f, y angles.  As a result the BDEs increase by about 10 kJ mol-1 in b-sheet and 40 kJ mol-1 in a-helical environments, respectively. In effect the aC-H BDEs can be "tuned" from about 345 to 400 kJ mol-1 by adjusting the local environment.  Some very significant effects of this are seen in the current literature on H transfer processes in enzyme mechanisms and in oxidative damage to proteins.  These observations are discussed in terms of the findings of the present study.- Return to Selected Publications


140. Oxidative Damage  to and by Cysteine in Proteins: an Ab Initio Study of the Radical Structures, C-H, S-H and C-C Bond Dissociation Energies, and Transition structures for H Abstraction by Thiyl Radicals

Abstract:  Ab initio computations (B3LYP/6-31G(D), coupled with isodesmic reactions) were used to predict bond dissociation energies (BDEs) of  aC-H (D(aC-H)) and other bonds of cysteine, both as free neutral amino acid (AH(Cys)) and as a residue in a model peptide (PH(Cys)).  The latter was intended to mimic the environment in proteins.  Transition structures were located for intermolecular and intramolecular H atom transfer to a thiyl radical (RS.) from a sulfhydryl group (RSH) or the aC-H bond.  The predicted BDEs, at 298 K,  in kJ mol-1 to an estimated accuracy of 10 kJ mol-1 for the fully optimized system are: AH(Cys), D(aC-H) = 322, D(bC-H) = 390,  D(aC-C)= 264,  D(S-H)=  373;  PH(Cys),  D(aC-H)= 346,  D(bC-H)= 392,  D(aC-C)= 287,  D(S-H)= 367.  In PH(Cys) with torsional angles constrained to simulate b-sheet and a-helical secondary structure,  rises to 359 and 376, respectively. Cystine in the peptide environment was modelled by replacing -SH by -SSCH3,  PH(CysSCH3), D(aC-H) = 330.  Enthalpies of activation for intermolecular H transfer to RS. were found to be low: from RSH, 12 kJ mol-1; from aC-H, about 25 kJ mol-1, the latter being consistent with reaction rates of the order of 105 M-1 s-1.  The enthalpic barrier for intramolecular H transfer from aC-H to -S. within a single cysteine residue is too high (83 - 111 kJ mol-1) for this to be a competitive process.- Return to Selected Publications


128. Toward Site Specificity of Oxidative Damage in Proteins: C-H and C-C Bond Dissociation Energies and the Reduction Potentials of the Radicals of Alanine, Serine, and Threonine - an Ab Initio Study

Abstract: High level ab initio computations were used to characterise the parent species and radicals for alanine, serine and threonine, both as free neutral amino acids (AH) and as residues in model peptides (PH) intended to mimic the mid chain environment in proteins. The ab initio energies were used in isodesmic reactions to predict bond dissociation energies (BDEs, ) at 298 K, in kJ mol-1 to an estimated accuracy of 10 kJ mol-1. For the fully optimized systems the values of are: AH(Gly), 331; AH(Ala), 317; AH(Ser), 327; AH(Thr), 328; PH(Gly), 348; PH(Ala), 344; PH(Ser), 348; PH(Thr), 356. All of the values are less than the BDE of a typical SH bond (370 kJ mol-1), as in cysteine or glutathione (GSH), a result that suggests that oxidative damage at the site will not be repaired efficiently by the mechanism of H donation from GSH. Values of in typical peptide conformations, such as -sheet and -helical secondary structure, were estimated by constraining the Ramachandran dihedral angles phi, and psi, to values typical of these structures. Thus values are estimated as: PH(Gly), 361; PH(Ala), 359; PH(Ser), 347; PH(Thr), 356 in the beta-sheet conformation, and: PH(Gly), 402; PH(Ala), 384; PH(Ser), 381; PH(Thr), 363 in the alpha-helix conformation. Hence, these residues are also expected to be susceptible to irreparable oxidative damage in beta-sheet structures, but Gly, Ala and Ser residues in alpha-helical regions should be less susceptible to damage and should be repairable by GSH. A consideration of reduction potentials calculated from the BDEs and entropies derived from the ab initio results leads to the same conclusions and indicates that certain radicals other than OH. that occur in cells (e.g. ROO.) may also cause oxidative damage to beta-sheet structures. Ab initio calculations were also done for the C-centered radicals formed by removal of H from the side chains. These showed that there is a marked increase in the ease of abstraction of this H in the series Ala, Ser,Thr. - Return to Selected Publications